People
Current Lab Members
Joseph Groenke (B.S./B.A. University of Michigan)
Research Assistant & Laboratory Coordinator (2017 - present)
Department of Biomedical Sciences, Heritage College of Osteopathic Medicine.
Email
Joe focuses on the technical development of biological and paleontological data through digital and mechanical preparation. He differentiates computed tomographic volumes used to characterize morphology, and he creates biologically realistic polygon products from those differentiations that are used to test developmental, functional, anatomical, and phylogenetic hypotheses. He is responsible for the organization, archiving, replication and distribution of lab data, and trains students to understand the importance of transparent, repeatable methodologies, data stewardship, and data sharing within the greater scientific community through the NSF-funded MorphoSource. His recent efforts have involved elucidating phenotypic aspects of growth, development, and pathology in model organisms (turkey, mice). He has also worked extensively with fossil material from the Late Cretaceous of Madagascar and Tanzania, as well as serving in logistical coordinating roles in six field seasons in Madagascar between 2005 and 2023. He has also been a field crew member for other work based in Eastern Africa and Western North America.
Selected Publications:
Krause, D. W., S. Hoffmann, J. R. Wible, E. C. Kirk, J. A. Schultz, W. v. Koenigswald, J. R. Groenke
, J. B. Rossie, P. M. O’Connor, E. R. Seiffert, E. R. Dumont, W. L. Holloway, R. R. Rogers, L. J. Rahantarisoa, A. D. Kemp, and H. Andriamialison. 2014.
First cranial remains of gondwanatherian mammal reveal remarkable mosaicism. Nature
515
:512-517.
O’Connor, P. M., D. W. Krause, N. J. Stevens, J. R. Groenke , R. D. E. MacPhee, and D. Kalthoff. 2019. A new mammal from the Upper Cretaceous (Turonian - Campanian) Galula Formation, southwestern Tanzania. Acta Paleontologica Polonica 64 (1) :65-84.
O’Connor, P. M., A. H. Turner, J. R. Groenke , R. R. Rogers, D. W. Krause, and L. J. Rahantarisoa. 2020 . Late Cretaceous bird from Madagascar reveals unique development of beaks. Nature 588 :272-276 doi.org/10.1038/s41586-020-2945-x .
J. R. Groenke & P.M. O’Connor. 2023 . Enhanced resolution at the mechanical-digital interface: A methodology to assist in preparation decision making and improved research and outreach outcomes. Society of Vertebrate Paleontology 83rd annual meeting, Cincinnati, OH, October 2023.
Joe Groenke- GoogleScholar Profile: Groenke GS
Chris Torres (B.S. North Carolina State University; Ph.D., University of Texas Austin)
National Science Foundation Postdoctoral Fellow (2021 - 2024)
Department of Biomedical Sciences, Heritage College of Osteopathic Medicine.
Email
Chris is an evolutionary morphologist focused on how the relationships between organisms and their environments change, how the shapes of those organisms change in response, and how those changes have shaped the diversity we see in the world today… and the diversity we don’t see anymore. In the O’Connor lab, Chris is investigating what the shape of the brain can tell us about how birds experience their worlds both today and in the distant past. Chris’ paleontological field work includes expeditions in the US, Southern Chile and Antarctica. He also really, really likes flamingos.
Selected Publications:
Torres, C. R.
, M. A. Norell, and J. A. Clarke. 2021
. Bird neurocranial and body mass evolution across the end-Cretaceous mass extinction: The avian brain shape left other dinosaurs behind. Science Advances
7
:eabg7099.
Torres, C. R. , M. A. Norell, and J. A. Clarke. 2021 . Estimating flight style of Early Eocene stem palaeognath bird Calciavis grandei (Lithornithidae). The Anatomical Record 303 :1035-1042.
Torres, C. R. and J. A. Clarke. 2018 . Nocturnal giants: evolution of the sensory ecology in elephant birds and other paleognaths inferred from digital brain reconstructions. Proceedings B 285 :20181540.
CR Torres , VL De Pietri, A Louchart and M van Tuinen. 2015 . New cranial material from the earliest filter feeding flamingo Harrisonavis croizeti (Aves, Phoenicopteridae) informs the evolution of the highly specialized filter feeding apparatus. Organisms, Evolution & Diversity s. Organisms, Evolution & Diversity 153 :609-618.
Chris Torres-GoogleScholar Profile: Torres GS
Belal Salem (B.S. Benha University, Egypt; M.S. Biological Sciences, Ohio University)
Current Ph.D. Student (2024 - present)
Graduate Program in Biological Sciences
Ohio Center for Ecology and Evolutionary Studies
Email
Belal is interested in the paleontology and comparative anatomy of northern African Late Cretaceous Archosaurs (including dinosaurs, crocodiles, pterosaurs, etc) with an emphasis on the Egyptian record. He acted as the field manager at the Mansoura University Vertebrate Paleontology center (MUVP) in Egypt from 2020 to 2022. Belal has been actively involved in roughly 28 field expeditions in Egypt collecting and curating a wide array of vertebrate fossils. Thus far, Belal has two publications that focus on new dinosaur remains from the Late Cretaceous of Egypt (see below). He also works at Benha University in Egypt as a teaching assistant. After receiving his PhD, he plans to continue in a research position and be the first faculty to study vertebrate paleontology in the history of Benha University.
Selected Publications:
Salem, Belal S.
, M. C. Lamanna, P. M. O'Connor, W.A. Thabet, S. El-Sayed, H.M. Sallam. 2023
. A peirosaurid crocodyliform from the Upper Cretaceous (Cenomanian) Bahariya Formation of the Bahariya Oasis, Western Desert, Egypt. Annual meeting for the Society of Vertebrate Paleontology, Program and Abstracts, pp. 378-379.
Salem, Belal S.
, M. C. Lamanna, P. M. O'Connor, G. M. El-Qot, F. A. Shaker, W. A., Thabet, S. El-Sayed, and H. M. Sallam. 2022
. First definitive record of Abelisauridae (Theropoda: Ceratosauria) from the Cretaceous Bahariya Formation, Bahariya Oasis, Western Desert of Egypt. Royal Soceity Open Science https://dx.doi.org/10.1098/rsos.220106 PDF available upon request
Salem, Belal S.
, P. M. O'Connor, E. Gorscak, S. El-Sayed, J. J. W. Sertich, E. Seiffert, and H. M. Sallam. 2021
. Dinosaur remains from the Upper Cretaceous (Campanian) of the Western Desert, Egypt. Cretaceous Research 123: https://doi.org/10.1016/j.cretres.2021.104783 PDF available upon request
Belal Salem-GoogleScholar Profile: Salem GS
Riley Sombathy (B.S. Troy University/M.Sc. Adelphi University)
Current Ph.D. Candidate (Fall 2021 - present)
Graduate Program in Biological Sciences
Ohio Center for Ecology and Evolutionary Studies
Email
Riley studies paleohistology with a focus on how the growth rate and body mass of vertebrates influence the development of bone tissue. His current research includes works on both extinct and extant theropods, using the clade as a system for investigating various factors in bone tissue development. He received his MS degree from Adelphi University in 2021 and has field experience across Wyoming.
Selected Publications:
Sombathy, R.
, M. D’Emic, and P. M. O’Connor. 2023
. Osteohistology of the theropod dinosaur Ceratosaurus
. Annual meeting for the Society of Vertebrate Paleontology, Program and Abstracts, p. 406.
Bakoli Rakotozafy (B.S./M.S., University of Antananarivo, Antananarivo, Madagascar)
Ph.D. Student (2018 - 2024)
Menton BEC (Bassins sedimentairies-Evolution-Conservation), University of Antananarivo, Madagascar
Bakoli completed her MS in 2016. Entitled Ontogenie de Mahajangasuchus insignis , Mesoeucrocodylia de la Formation de Maevarano, Bassin de Mahajanga. In early June 2024, she defended her dissertation entitled, Écomorphologie des crocodyliformes de la Formation Maevarano du Crétacé supérieur du Bassin de Mahajanga, Nord-ouest de Madagascar.
Bakoli has conducted field and laboratory research with MBP teams since 2015, in addition to working with our group and the faculty at University of Antananarivo (UA) on collections/facilities efforts. Bakoli completed (November 2022 - December 2023) an extended trainingship at the Denver Museum of Nature & Science. In addition to assisting with teaching at the University of Antananarivo, Bakoli will oversee the fossil collection facility developed at the University.
Selected Publications:
Rakotozafy, B.
, J. J. W. Sertich, and P. M. O’Connor. 2023
. Cranial ontogeny in the notosuchian crocodyliform Miadanasuchus oblita
from the Maevarano Formation, northwestern Madagascar. Annual meeting for the Society of Vertebrate Paleontology, Program and Abstracts, pp. 358-359.
Rakotozafy, B.
, L. Ranivoharimanana, J. J. W. Sertich, A. Rasoamiaramanana and P. M. O’Connor. 2024
. Late Cretaceous Notosuchians of the Maevarano Formation, Mahajanga Basin: An Ecomorphological Perspective. Madamines, volume 1:1-53, 2024 (ISSN 2220-0681). PDF Available Here
Eric Lund (B.S./M.Sc. University of Utah)
Current Ph.D. Student (2014 - present)
Graduate Program in Biological Sciences and Geological Sciences
Individual Interdisciplinary Program, Ohio Center for Ecology and Evolutionary Studies
Lab Coordinator (2011-2016), Department of Biomedical Sciences, Ohio University
Current Position (2017 - present): Paleontology Lab Manager & Paleontology Lab Coordinator, North Carolina Museum of Natural Science
Eric is interested in the paleobiology and functional morphology of ceratopsian (horned) dinosaurs and faunal evolution in the Western Interior Basin during the Cretaceous Period. He received his MS degree from the University of Utah in 2010. He has had the opportunity to conduct paleontological field work in Utah, Wyoming, and around the world including Mexico, Tanzania and Madagascar. Eric first moved to Ohio University in late 2011 to work as a laboratory coordinator with the O'Connor lab. He later matriculated as a PhD student in the IIP, focusing his research around ceratopsid evolutionary morphology and functional anatomy. Eric is currently completing his Ph.D. while also serving as the Fossil Conservation Laboratory Manager at the North Carolina Museum of Natural Sciences working with Dr. Lindsay Zanno.
Selected Publications:
Lund, E. K.
, P. M. O’Connor, M. A. Loewen, and Z. A. Jinnah. 2016
. A New Centrosaurine Ceratopsid, Machairoceratops cronusi
gen et sp. nov., from the upper sand member of the Wahweap Formation (Middle Campanian), Southern Utah. PLoS ONE
11(5)
:e0154403. doi:10.1371/journal.pone.0154403. PDF available upon request
Sampson, S. D., E. K. Lund , M. A. Loewen, A. A. Farke, and K. E. Clayton. 2013 . A remarkable short snouted horned dinosaur from the Late Cretaceous (late Campanian) of southern Laramidia. Proceedings of the Royal Society B: Biological Sciences 280 (1766):20131186.
Former Graduate Students and Postdoctoral Researchers
Ryan Felice (B.S., Tulane University)
Former PhD Student, Completed Ph.D. 2015
Graduate Program in Biological Sciences (2009 - 2015)
Ohio Center for Ecology and Evolutionary Studies
Current Position: Lecturer in Human Anatomy, Centre for Integrative Anatomy, Department of Cell and Developmental Biology, University College London
Ryan is interested in the paleobiology, evolution, and functional morphology. His dissertation was entitled: Evolution and Integration of Avian Caudal Skeletal Morphology . Other research efforts included field work in Wyoming, Utah and Tanzania and a study of synapsid locomotion. After completing his PhD in 2015, Ryan moved to the UK to complete a postdoctoral research fellowship in the lab group of Dr. Anjali Goswami at UC London/Natural History Museum-London. Ryan and Pat currently collaborate on analyses of cranial shape dispartiy and evolution in birds and crocodyliforms.
Selected Publications from PhD:
Felice, R.N., and P. M. O'Connor. 2016 Function drives caudal skeletal variation in sexually dimorphic passeriforms. Journal of Avian Biology: 47:371-377. doi: 10.1111/jav.00801
Felice, R.N. 2014 (early view). Coevolution of Caudal Skeleton and Tail Feathers in Birds. Journal of Morphology. doi: 10.1002/jmor.20321
Felice, R. N. , and P. M. O’Connor. 2014 . Ecology and caudal skeleton morphology in birds: The convergent evolution of pygostyle shape in underwater foraging taxa. PLoS One 9(2): e89737. doi:10.1371/journal.pone.0089737.
Sam Gutherz (B.S. Georgetown University)
Graduate Program in Biological Sciences (2016 - 2022)
Ohio Center for Ecology and Evolutionary Studies
Current Position (2022-present): Instructor of Anatomical Sciences, Midwestern University-Chicago College of Osteopathic Medicine
Research Interests:
Sam is interested in the interactions between pulmonary tissue and the skeleton in birds. By studying the development of postcranial pneumaticity in living birds, he hopes to address questions regarding the evolution of the respiratory system in a range of archosaurs. Thus far Sam has published the first and second of his dissertation chapters, one focused on cuckoos, with the other on Accipitrimorphae. His past research includes neurobiology, paleopathology, and field research in North Dakota and Montana.
Selected Publications from PhD:
Gutherz, S. B.
, J. R. Groenke, J. J. W. Sertich, S. H. Burch, and P. M. O'Connor. 2020
. Paleopathology in a nearly complete skeleton of Majungasaurus crenatissimus
(Theropoda: Abelisauridae). Cretaceous Research 115:e104553 https://doi.org/10.1016/j.cretres.2020.104553 PDF available upon request
Gutherz, S. B.
and P. M. O'Connor. 2021
. Postcranial skeletal pneumaticity in Cuclidae. Zoology https://doi.org/10.1016/j.zool.2021.125907 PDF available upon request
Gutherz, S. B.
and P. M. O'Connor. 2022
. Postcranial skeletal pneumaticity in non-aquatic neoavians: Insights from accipitrimorphae. J Anatomy https://onlinelibrary.wiley.com/doi/full/10.1111/joa.13742 PDF available upon request
Eric Gorscak (B.S. Ohio University)
Former Ph.D. Student, Completed Ph.D. 2016
Graduate Program in Biological Sciences (2010-2016)
Ohio Center for Ecology and Evolutionary Studies
Current Position: Assistant Professor in Anatomical Sciences, Midwestern University-Chicago College of Osteopathic Medicine
EmailResearch Interests:
Eric is interested phylogenetics, biogeography, and the evolution of African dinosaurs and other vertebrates during the Cretaceous Period. His dissertation was entitled: Descriptive and Comparative Morphology of African Titanosaurian Sauropods: New Information on the Evolution of Cretaceous African Continental Faunas
. After completing his PhD in 2016, Eric undertook a postdoctoral research fellowship at the Field Museum of Natural History (Chicago) with Dr. Peter Makovicky. A portion of his research falls under the larger Rukwa Rift Basin Project
of southwestern Tanzania. During his PhD, Eric participating in number field expeditions, including those associated with ongoing field research in Tanzania, Antarctica, Utah, and Madagascar.
Selected Publications from PhD:
Gorscak, E. , P. M. O’Connor, N. J. Stevens, and E. M. Roberts. 2017 . The second titanosaurian (Dinosauria: Sauropoda) from the middle Cretaceous Galula Formation, southwestern Tanzania with remarks on African titanosaurian diversity. Journal of Vertebrate Paleontology 37: http://dx.doi.org/10.1080/02724634.2017.1343250.PDF available upon request
Gorscak, E. and P. M. O’Connor. 2016 . Time-calibrated models support congruency between Cretaceous rifting and titanosaurian evolutionary history. Biology Letters 12 :20151047 doi.org/10/1098/rsbl.2015.1047. PDF available upon request
Gorscak, E. , P. M. O'Connor, N. J. Stevens, and E. M. Roberts. 2014 . The basal titanosaurian Rukwatitan bisepultus (Dinosauria, Sauropoda) from the middle Cretaceous Galula Formation, Rukwa Rift Basin, southwestern Tanzania. Journal of Vertebrate Paleontology 34:1133–1154. PDF available upon request
Waymon Lewis Holloway (B.S. Ohio University/M.S. Marshall University)
Former Ph.D. Student, Completed Ph.D. 2018
Graduate Program in Biological Sciences (2013 - 2018)
Ohio Center for Ecology and Evolutionary Studies
Former Position: Postdoctoral Instructor of Clinical Gross Anatomy, Department of Anatomical Sciences, Renaissance School of Medicine, Stony Brook University, Stony Brook, NY, USA (2018 - 2021).
Current Position: Assitant Professor of Biology, Baldwin Wallage University (2021 - present).
Waymon is interested in the functional anatomy of the head and sensory system evolution of archosaurs. His dissertation was entitled: Comparative Cranial Ecomorphology and Functional Morphology of Semiaquatic Faunivorous Crurotarsans . His past research has included invertebrate and microfloral surveys and a digital phytosaur endocast reconstruction, the latter as part of his MS project at Marshall University.
Selected publications from graduate school:
Holloway, W. L., K. M. Claeson, H. M. Sallam, J. J. W. Sertich, and P. M. O’Connor. 2017 . A new species of the neopterygian fish Enchodus from the Duwi Formation, Campanian, Late Cretaceous, Western Desert, central Egyptr. Acta Palaeontologica Polonica 62 :603-611. doi:https://doi.org/10.4202/app.00331.2016.
Krause, D. W., S. Hoffmann, J. R. Wible, E. C. Kirk, J. A. Schultz, W. v. Koenigswald, J. R. Groenke, J. B. Rossie, P. M. O’Connor, E. R. Seiffert, E. R. Dumont, W. L. Holloway, R. R. Rogers, L. J. Rahantarisoa, A. D. Kemp, and H. Andriamialison. 2014 . First cranial remains of gondwanatherian mammal reveal remarkable mosaicism. Nature 515 :512-517.
Holloway, W.L., K.M. Claeson, and F.R. O’Keefe. 2013 . A virtual phytosaur endocast and its implications for sensory system evolution in archosaurs. Journal of Vertebrate Paleontology 33 :848-857.
Nirina (Liva) Ratsimbaholison
Former Ph.D. Student, Completed Ph.D. 2016
Biological Anthropology and Paleontology, University of Antananarivo, Madagascar
Affiliated Student, Ohio Center for Ecology and Evolutionary Studies
Liva completed a Ph.D. in 2016 entitled: Ontogenetic Trends in the Craniomandibular Skeleton of Abelisauridae. He has conducted field research along with the MBP field teams since 2005. Since completing his Ph.D. he has returned to Madagascar, completed a research visit in India, and is continuing to work on publishing dissertation chapters with P. O'Connor and T. Hieronymus.
DEA [Masters Degree] in Paleontology, Université d’Antananarivo, Madagascar. Project Title: Ontogenie Craniofaciale de Majungasaurus crenatissimus (Theropode: Abelisauridae) du Cretace Superieur de Berivotra-Mahajanga. Thesis Defended: April, 2008.
Project Title: Ontogenetic variability of cranio-mandidular system in Majungasaurus crenatissimus .
Selected Publications from PhD:
Ratsimbaholison. N. O.
, R. N. Felice, and P. M. O’Connor. 2016
. Ontogenetic changes in the craniomandibular skeleton of Majungasaurus crenatissimus
from the Late Cretaceous of Madagascar. Acta Palaeontologica Polonica
61
:281-292 (2016)
. PDF available upon request
Kerin M. Claeson, Ph.D.
Postdoctoral Researcher April 2010-June 2012
Department of Biomedical Sciences, Anatomy Instructor
Ohio Center for Ecology and Evolutionary Studies
Current Position: Professor of Anatomy, Philadelphia College of Osteopathic Medicine
Email
Kerin’s research combines the disciplines of paleontology, comparative anatomy, and development to formulate and test phylogenetic hypotheses of interrelationships of several groups of fishes, and occasionally other vertebrates. She makes extensive use of worldwide museum collections but has also had the opportunity to conduct field work in Argentina, Pennsylvania, Texas, and Wyoming. She received her PhD in Geological Sciences from The University of Texas at Austin in 2010 and her Masters in Organismic and Evolutionary Biology from the University of Massachusetts at Amherst in 2005. Kerin started a faculty position at the Philadelphia College of Osteopathic Medicine in July 2012. She has been promoted to associate, and now (2021) full professor, where she continues her research on fishes while also playing a significant role in educating future physicians.
Ph.D. Department of Biological Sciences (2004 - 2009)
Ohio Center for Ecology and Evolutionary Studies
Project Title: Evolution of the avian wing skeleton: Integrating whole bone morphology, cross-sectional geometry, histology, and mechanical testing for evaluating the relationship between form and function in the forelimb of birds.
Current Position: Professor of Anatomy, Midwestern University (Glendale, AZ).
Selected Publications from PhD:
Simons, E. L. R. and P. M. O’Connor. Bone laminarity in the avian forelimb skeleton and its relationship to flight mode: Testing Functional Interpretations.
Anatomical Record
295:386-396 (2012)
.
Simons, E. L. R. , T. L. Hieronymus, and P. M. O’Connor. Cross sectional geometry of the forelimb skeleton and flight mode in pelecaniform birds . J Morphology 272(8):958-971 (2011) .